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List of doctoral-level courses


Registration for compulsory comprehensives (BIOC 701, 702, 703)

All students entering or registered in the Ph.D. program must register for the comprehensive courses (see table below). Registration for these courses is not initiated until instructed by the Student Affairs Officer (Christine Laberge).

Questions concerning registration for comprehensives should be directed to the Student Affairs Officer. Please refer to the guidelines for Research Seminar 1, Ph.D. Thesis Proposal, and Research Seminar 2.


Required Courses (3 credits)

A minimum attendance at 70% of seminars, including Junior and Senior Seminars, is required to pass BIOC 696

BIOC 696D1 Seminars in Biochemistry 1.5
BIOC 696D2 Seminars in Biochemistry 1.5
BIOC 701 Research Seminar 1 0
BIOC 702 Ph.D. Thesis Proposal 0
BIOC 703 Research Seminar 2 0


Students must complete BIOC 701 in the third term after admission to the program, BIOC 702 in the 5th or 6th term, and BIOC 703 approximately 6 months prior to submissin of the Ph.D. thesis.

Complementary Courses (6 credits)

At least one course from the courses listed below plus additional credits to a minimum of 6 total complementary course credits of 500-level or higher courses in the biomedical and allied sciences, chosen in consultation with the research director.

BIOC 570 Biochemistry of Lipoproteins 3
BIOC 600 Adv Strat in Genetics&Genomics 3
BIOC 603 Genomics and Gene Expression 3
BIOC 604 Macromolecular Structure 3
BIOC 605 Protein Biology & Proteomics 3
EXMD 615 Essentials of Glycobiology 3
EXMD 635D1 Experimental/Clinical Oncology 3
EXMD 635D2 Experimental/Clinical Oncology 3


The Graduate Advisory Committee may stipulate additional course work depending on the background of the candidate. BIOC 450 (Protein Structure and Function) and BIOC 454 (Nucleic Acids) are additional requirements for those who have not previously completed equivalent courses in their prior training.

Key dates